一、 简介

杨贤鹏，男，理学博士，副教授，硕士生导师。邮箱：yangxp2006@sdnu.edu.cn

工作经历:

2023/12-至今 山东师范大学生命科学学院 副教授，硕士生导师

2019/07-2023/12 山东师范大学生命科学学院 讲师，硕士生导师

2017/07-2019/06 中国科学院武汉植物园 博士后

教育经历:

2013/09-2017/06 中国科学院武汉植物园 植物学 博士

2010/09-2013/06 中国科学院近代物理研究所 生物物理学 硕士

2006/09-2010/06 吉林大学 生物技术 学士

二、 研究方向

研究方向一：藜麦木栓质的合成调控分子机制解析

木栓质是植物根系的疏水性脂质屏障结构，在植物的水分和营养物质吸收过程中起重要作用。利用耐盐作物藜麦为研究材料，深入探讨木栓质的合成以及响应环境非生物和营养胁迫的分子调控机制。

研究方向二：小麦和拟南芥表皮蜡质的合成调控机制解析

蜡质是覆盖在植物最外层的疏水性脂质结构，通过限制水分的非气孔性散失进而提高植物对非生物胁迫的耐受性。利用小麦和拟南芥为研究材料，深入解析蜡质合成响应干旱和盐胁迫的分子调控机制。

三、 承担项目

1. 国家自然科学基金面上项目：TaCER9负调控小麦蜡质合成和耐旱性的分子机制研究（32670363），2027-2030，50万，在研；

2. 国家自然科学基金青年基金项目：乙酰辅酶A羧化酶ACC1参与拟南芥蜡质合成响应盐胁迫的功能解析（32100207），2022-2024，30万，结题；

3. 山东省自然科学基金青年基金项目：长链脂肪醇氧化酶FAO3和FAO4b参与拟南芥角质组分α,ω二元羧酸合成的功能解析（ZR2020QC018），2021-2023，15万，结题；

4. 中国博士后科学基金面上项目二等资助：长链脂肪醇氧化酶FAO3参与表皮蜡质降解的功能解析（2018M642961），2018-2019，5万，结题。

四、 教学科研成果（论文、专著、专利等）

研究论文 

1. Wang Z, Huang H, Lü S, Wang P, Yang X*, Ma C* (2026) The MYB41-MYB74 module and VOZ1 transcription factor synergistically regulate aliphatic suberin monomer biosynthesis in Chenopodium quinoa. The Plant Journal 126: e70914

2. Wang Z, Liu Y, Huang H, Zheng Z, Lü S, Yang X*, Ma C* (2025) Functional identification of two Glycerol-3-phosphate Acyltransferase5 homologs from Chenopodium quinoa. Plant Science 350: 112313

3. Li S^#, Zhang X^#, Huang H^#, Yin M, Jenks MA, Kosma DK, Yang P, Yang X*, Zhao H*, Lü S* (2025) Deciphering the core shunt mechanism in Arabidopsis cuticular wax biosynthesis and its role in plant environmental adaptation. Nature Plants 11: 165-175

4. Yang X, Huang H, Wang Z, Haslam TM, Kunst L, Wang P, Zhao H, Lü S, Ma C (2025) Acetyl-CoA Carboxylase1 influences ECERIFERUM2 activity to mediate the synthesis of very-long-chain fatty acid past C28. Plant Physiology 197 (2): kiae253

5. Li L^#, Yang X^#, Huang H, Zhu C, Xing M, Yang K, Nie X, Fu J, Wang M, Liang Z, Lü S, Qian W, Tran LP*, Yin X*, Li W* (2025) The small nucleolar RNA NON-CODING RNA 1 negatively regulates drought tolerance in Arabidopsis thaliana. The Plant Journal 123: e70428

6. Huang H, Wang Y, Yang P, Zhao H, Jenks MA, Lü S*, Yang X* (2024) The Arabidopsis cytochrome P450 enzyme CYP96A4 is involved in the wound-induced biosynthesis of cuticular wax and cutin monomers. The Plant Journal 118: 1619-1634

7. Huang H^#, Yang X^#, Zheng M, Chen Z, Yang Z, Wu P, Jenks MA, Wang G, Feng T, Liu L, Yang P, Lü S*, Zhao H* (2023) An ancestral role for 3-KETOACYL-COA SYNTHASE3 as a negative regulator of plant cuticular wax synthesis. The Plant Cell 35: 2251-2270

8. Li S^#, Yang X^#, Huang H, Qiao R, Jenks MA, Zhao H, Lü S (2022) Arabidopsis ACYL-ACTIVATING ENZYME 9 (AAE9) encoding an isobutyl-CoA synthetase is a key factor connecting branched-chain amino acid catabolism with iso-branched wax biosynthesis. New Phytologist 233: 2458-2470

9. Tariq F^#, Zhao S^#, Ahmad N, Wang P, Shao Q, Ma C*, Yang X* (2022) Overexpression of β-Ketoacyl CoA Synthase 2B.1 from Chenopodium quinoa promotes suberin monomers' production and salt tolerance in Arabidopsis thaliana. International Journal of Molecular Sciences, 23(21), 13204.

10. Yang X^#, Cui L^#, Li S, Ma C, Kosma DK, Zhao H*, Lü S* (2022) Fatty alcohol oxidase 3 (FAO3) and FAO4b connect the alcohol- and alkane-forming pathways in Arabidopsis stem wax biosynthesis. Journal of Experimental Botany 73: 3018-3029

11. Yang X, Feng T, Li S, Zhao H, Zhao S, Ma C, Jenks MA, Lü S (2020) CER16 inhibits post-transcriptional gene silencing of CER3 to regulate alkane biosynthesis. Plant Physiology 182: 1211-1221

12. Yang X^#, Wang Z^#, Feng T, Li J, Huang L, Yang B, Zhao H, Jenks MA, Yang P*, Lü S* (2018) Evolutionarily conserved function of the sacred lotus (Nelumbo nucifera Gaertn.) CER2-LIKE family in very-long-chain fatty acid elongation. Planta 248: 715-727

13. Yang X^#, Zhao H^#, Kosma DK, Tomasi P, Dyer JM, Li R, Liu X, Wang Z, Parsons EP, Jenks MA, Lü S (2017) The acyl desaturase CER17 is involved in producing wax unsaturated primary alcohols and cutin monomers. Plant Physiology 173: 1109-1124

合作论文

1. Li L, Yang X, Huang H, Zhu C, Xing M, Yang K, Nie X, Fu J, Wang M, Liang Z, Feng X, Zhao J, Li X, Lü S, Shi Y, Tran LP, Yin X, Li W (2026) Small nucleolar RNA HIDDEN TREASURE 2 reduces drought tolerance via multiple pathways in Arabidopsis. The Plant Journal 125: e70718

2. Li X, Su T, Wang X, Liu Y, Ge J, Huo P, Zhao Y, Wang T, Yu H, Duan M, Jia Y, Yang X, Wang P, Gong Q, Liu J, Ma C (2025) Blocking constitutive autophagy rescues the loss of acquired heat resistance in Arabidopsis fes1a. New Phytologist 245: 2569-2583

3. Wang H, Lu Z, Xu Y, Zhang J, Han L, Chai M, Wang ZY, Yang X, Lü S, Tong J, Xiao L, Wen J, Mysore KS, Zhou C (2023) Roles of very long-chain fatty acids in compound leaf patterning in Medicago truncatula. Plant Physiology 191: 1751-1770

4. Su Y, Feng T, Liu CB, Huang H, Wang YL, Fu X, Han ML, Zhang X, Huang X, Wu JC, Song T, Shen H, Yang X, Xu L, Lü S, Chao DY (2023) The evolutionary innovation of root suberin lamellae contributed to the rise of seed plants. Nature Plants 9: 1968-1977

5. Huang H, Yang X, Zheng M, Lu S, Zhao H (2023) Fine-tuning the activities of beta-KETOACYL-COA SYNTHASE 3 (KCS3) and KCS12 in Arabidopsis is essential for maintaining cuticle integrity. Journal of Experimental Botany 74: 6575-6587

6. Yang Q, Yang X, Wang L, Zheng B, Cai Y, Ogutu CO, Zhao L, Peng Q, Liao L, Zhao Y, Zhou H, Han Y (2022) Two R2R3-MYB genes cooperatively control trichome development and cuticular wax biosynthesis in Prunus persica. New Phytologist 234: 179-196

7. Wang Y, Yang X, Chen Z, Zhang J, Si K, Xu R, He Y, Zhu F, Cheng Y (2022) Function and transcriptional regulation of CsKCS20 in the elongation of very-long-chain fatty acids and wax biosynthesis in Citrus sinensis flavedo. Horticulture Research 9, uhab027.

8. Huang H, Ayaz A, Zheng M, Yang X, Zaman W, Zhao H, Lü S (2022) Arabidopsis KCS5 and KCS6 Play Redundant Roles in Wax Synthesis. International Journal of Molecular Sciences 23(8), 4450

9. Zhao Z, Yang X, Lü S, Fan J, Opiyo S, Yang P, Mangold J, Mackey D, Xia Y (2020) Deciphering the Novel Role of AtMIN7 in Cuticle Formation and Defense against the Bacterial Pathogen Infection. International Journal of Molecular Sciences 21(15), 5547

10. Zhang P, Wang R, Yang X, Ju Q, Li W, Lü S, Tran LP, Xu J (2020) The R2R3-MYB transcription factor AtMYB49 modulates salt tolerance in Arabidopsis by modulating the cuticle formation and antioxidant defence. Plant, Cell and Environment 43: 1925-1943

11. Tang J, Yang X, Xiao C, Li J, Chen Y, Li R, Li S, Lü S, Hu H (2020) GDSL lipase occluded stomatal pore 1 is required for wax biosynthesis and stomatal cuticular ledge formation. New Phytologist 228: 1880-1896

12. Wang X, Zhou C, Yang X, Miao D, Zhang Y (2015) De Novo Transcriptome Analysis of Warburgia ugandensis to Identify Genes Involved in Terpenoids and Unsaturated Fatty Acids Biosynthesis. PLoS One 10: e0135724

五、 联系方式

山东师范大学（长清湖校区）博物楼A座308实验室